Biodiversity Data Journal Biodiversity Data Journal Biodiversity Data Journal BDJ 1314-2836 1314-2828 Pensoft Publishers Biodiversity Data Journal 10.3897/BDJ.2.e1165 3505 Taxonomic paper Nemata Faunistics & Distribution Catalogues and Checklists Nematodes from terrestrial and freshwater habitats in the Arctic Holovachov Oleksandr Dr oleksandr.holovachov@nrm.se † Swedish Museum of Natural History, Department of Zoology, Stockholm, Sweden

Corresponding author: Oleksandr Holovachov (oleksandr.holovachov@nrm.se).

Academic editor: Vlada Peneva.

 2014 19 08 2014 2 e1165 04 07 2014 14 08 2014 Oleksandr Holovachov This is an open access article distributed under the terms of the Creative Commons Attribution License 4.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.

We present an updated list of terrestrial and freshwater nematodes from all regions of the Arctic, for which records of properly identified nematode species are available: Svalbard, Jan Mayen, Iceland, Greenland, Nunavut, Northwest territories, Alaska, Lena River estuary, Taymyr and Severnaya Zemlya and Novaya Zemlya. The list includes 391 species belonging to 146 genera, 54 families and 10 orders of the phylum Nematoda.

 Alaska Arctic distribution fauna freshwater Greenland Iceland Jan Mayen Lena River Northwest territories Novaya Zemlya Nunavut Severnaya Zemlya Svalbard Taymyr terrestrial. "Taxonomy and distribution of free-living nematodes of the order Plectida in Sweden" and "Systematics of Swedish free-living nematodes of the orders Desmodorida and Araeolaimida" / Swedish Taxonomy Initiative Introduction

Nematodes are one of the most numerous and abundant multicellular organisms on the planet in general and in the Arctic in particular. There are over 70 research papers published which include data on the fauna and distribution of nematodes in the region. Few faunistic overviews of the Arctic nematodes exist in the literature. The "Catalogue of free-living soil and fresh-water nematodes of Arctic and Subarctic" (Kuzmin and Gagarin 1990) includes a large number of subarctic nematodes and is normally inaccessible. Subsequent publication by Gagarin (2001b) focuses only on the fresh-water nematodes from the Russian Arctic and Subarctic, while Coulson and Refseth (2004) give a list of terrestrial and freshwater nematodes from Svalbard and Jan Mayen. The present paper aims to provide a comprehensive overview of all findings of nematodes in the entire Arctic region between the years of 1883 and 2013, with all species names adjusted according to the most recent classification and nomenclature.

 Materials and methods

The species list has been compiled based on literature data and refers to the area of the Arctic (Fig. 1) as defined by the Circumpolar Arctic Vegetation Map (CAVM Team 2003) and Meltofte et al. (2013). The list below includes all species found in the Arctic, and specifies the regions where each species was found, as well as all relevant references. Current taxonomic status and systematic position of all species was updated according to the most recent views on nematode systematics. Section on nomenclature includes those synonymous names, and sometimes mistyped names, which were used in the original publications on the Arctic nematodes; it does not include any other existing synonyms or combinations. Asterisks indicate those new species names, which were originally proposed based on populations described from the Arctic.

Taxa identified in the literature to the genus level only were not considered in the review for the following reasons: 1) It is impossible to make sure how many actual species were considered by the author – just one, or if s/he had treated multiple species together in the same unit. 2) Taxa identified only to the genus level have limited usefulness when analyzing distribution, endemism and other aspects of biogeography. Nematode systematics is continuously improving. Menzel (1920) described Dorylaimus sp. from Spitzbergen (Svalbard). The context of the genus Dorylaimus had changed considerably since 1920 and now includes members of at least families Dorylaimidae, Qudsianematidae, Aporcelaimidae and Nordiidae, making it practically impossible to use Menzel's record in modern faunistic studies. Even genus-level identifications done more recently face similar problems. Species that are now considered to belong to the same genus may in future, when more data becomes available, be split into different, sometimes distantly related genera. It would thus be impossible to know in the future what author of the paper meant when he identified his material.

General classification and classification of the orders Chromadorida, Desmodorida and Araeolaimida follows De Ley and Blaxter (2004) and Holovachov and Esquivel (2012). Classification of the order Enoplida follows that of Smol et al. (2014) and Andrássy (2002); Triplonchida – Holovachov and Shoshin (2014); Dorylaimida – Andrássy (2009); Mononchida – Ahmad and Jairajpuri (2010); Monhysterida – Fonseca and Decraemer (2008); Plectida – Holovachov (2014); Rhabditida – Holovachov et al. (2009), Geraert (2008), Geraert (2011), Sudhaus (2011), Sudhaus and Fürst von Lieven (2003).

 Checklists Updated list of nematode species found in the Arctic Nematoda ENOPLIDA Ironidae Ironus tenuicaudatus de Man, 1876 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990).

 Alaimidae Alaimus arcuatus Thorne, 1939 Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Alaimus depressus Loof, 1971* Notes

Svalbard (Loof 1971).

 Alaimus parvus Thorne, 1939 Notes

Svalbard (Loof 1971).

 Alaimus primitivus de Man, 1880 Notes

Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Alaimus proximus Thorne, 1939 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Alaimus similis Thorne, 1939 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Cosalaimus striatus (Loof, 1964) Alaimus striatus Loof, 1964 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Amphidelus elegans (de Man, 1921) Alaimus elegans de Man, 1921 Notes

Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Paramphidelus dolichurus (de Man, 1876) Amphidelus dolichurus (de Man, 1876) Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Paramphidelus paludicola Gagarin, 1991* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991b, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Paramphidelus paramonovi (Eliashvili, 1971) Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Paramphidelus pusillus (Thorne, 1939) Amphidelus pusillus Thorne, 1939 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Rhabdolaimidae Rhabdolaimus terrestris de Man, 1880 Notes

Svalbard (Loof 1971); Nunavut, Canada (Mulvey 1969c); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Trefusiidae Tripylina arenicola (de Man, 1880) Trischistoma arenicola (de Man, 1880) Notes

Nunavut, Canada (Mulvey 1969c).

 Trischistoma monohystera (de Man, 1880) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 TRIPLONCHIDA Tobrilidae Tobrilus aberrans (W. Schneider, 1925) Notes

Northwest territories, Canada (Ebsary 1982).

 Tobrilus affinis Gagarin, 1996* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b).

 Tobrilus brevisetosus (W. Schneider, 1925) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Tobrilus gracilis (Bastian, 1865) Notes

Nunavut, Canada (Mulvey 1969c); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Gagarin and Kuzmin 1983, Kuzmin 1972, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Tobrilus helveticus (Hofmänner in Hofmänner & Menzel, 1914) Trilobus pseudallophysis Micoletzky 1925 Notes

Greenland (Ditlevsen 1927); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Kuzmin and Gagarin 1990).

 Tobrilus modestus Gagarin, 1996* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b).

 Tobrilus parvus Gagarin, 1991* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991a, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Tobrilus tripylis Gagarin, 1991* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991a, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Brevitobrilus stefanskii (Micoletzky, 1925) Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1999a, Gagarin 2001b).

 Epitobrilus allophysis (Steiner, 1919) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Eutobrilus affectiosus Shoshin, 1988 Eutobrilus antarcticus sensu Gagarin, 1993, 2001; Raritobrilus antarcticus sensu Gagarin 1990, 1991 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991a, Gagarin 1993, Gagarin 2001b, Gagarin 2001a, Gagarin 2009).

 Eutobrilus altherri (Andrássy, 1953) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b).

 Eutobrilus andrassyi (Altherr, 1963) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2009, Kuzmin and Gagarin 1990).

 Eutobrilus grandipapillatus (Brakenhoff, 1914) Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Eutobrilus selengaensis (Tsalolikhin, 1977) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b).

 Neotobrilus longus (Leidy, 1852) Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Kuzmin and Gagarin 1990).

 Paratrilobus grandipapilloides Micoletzky, 1922 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Paratrilobus rapis Gagarin, 1991* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991a, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Peritobrilus arcticus (Gagarin, 1991) Eutobrilus arcticus Gagarin, 1991* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991a, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Peritobrilus vipriensis Gagarin, 1999* Notes

Lena River estuary, Russia (Gagarin 1999b, Gagarin 2001b).

 Quasibrilus strenuus (Gagarin, 1991) Eutobrilus strenuus Gagarin, 1991* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991a, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Raritobrilus steineri (Micoletzky, 1925) Eutobrilus steineri (Micoletzky, 1925); Tobrilus steineri Micoletzky, 1925 Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Kuzmin 1986, Kuzmin and Gagarin 1990).

 Semitobrilus ebsaryi Tsalolikhin, 2000* Tobrilus longicaudatus sensu Ebsary, 1982 Notes

Northwest territories, Canada (Ebsary 1982).

 Semitobrilus longicaudatus (Schneider, 1923) Tobrilus longicaudatus (Schneider, 1923) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Semitobrilus parapellucidus (Ebsary, 1982) Tobrilus parapellucidus Ebsary, 1982* Notes

Northwest territories, Canada (Ebsary 1982); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Semitobrilus gagarini (Ebsary, 1982) Notes

Lena River estuary, Russia (Gagarin 2001b).

 Tripylidae Tripyla affinis de Man, 1880 Notes

Nunavut, Canada (Mulvey 1969c); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Tripyla dubia Gagarin, 1997* Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Tripyla filicaudata de Man, 1880 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Tripyla glomerans Bastian, 1865 Tripyla papillata Bütschli, 1873 Notes

Svalbard (Loof 1971); Greenland (Ditlevsen 1927); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Kuzmin 1972, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Tripyla infia Brzeski & Winiszewska-Slipinska, 1993 Tripyla filipjevi Altherr in Altherr & Delamare Deboutteville, 1972 Notes

Svalbard (Janiec 1997); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 1996, Gagarin 2001b, Gagarin 2001a).

 Tripyla magna Altherr in Altherr & Delamare Deboutteville, 1972 Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 1996, Gagarin 2001b, Gagarin 2001a).

 Tripyla setifera Bütschli, 1873 Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Prismatolaimidae Prismatolaimus dolichurus de Man, 1880 Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Greenland (Ditlevsen 1927); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Prismatolaimus intermedius (Bütschli, 1873) Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Nunavut, Canada (Mulvey 1969c); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Prismatolaimus primitivus Loof, 1971* Notes

Svalbard (Loof 1971).

 Prismatolaimus stenolaimoides Loof, 1971* Notes

Svalbard (Loof 1971).

 Prismatolaimus tareya Gagarin & Kuzmin, 1972* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1993, Gagarin and Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990).

 Prismatolaimus verrucosus Hirschmann, 1952 Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Bastianiidae Bastiania gracilis de Man, 1876 Notes

Svalbard (Loof 1971); Nunavut, Canada (Mulvey 1969c); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Bastiania parexilis De Coninck, 1935 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Odontolaimidae Odontolaimus chlorurus de Man, 1880 Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Diphtherophoridae Diphtherophora communis de Man, 1880 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Diphtherophora minuta Ivanova, 1958 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Diphtherophora obesa Thorne, 1939 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Diphtherophora perplexans (Cobb, 1913) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Tylolaimophorus bulgaricus (Andrássy, 1958) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Tylolaimophorus cylindricus (Cobb, 1920) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Trichodoridae Trichodorus californicus Allen, 1957 Notes

Alaska (Bernard 1992).

 Trichodorus carlingi Bernard, 1992* Notes

Alaska (Bernard 1992).

 DORYLAIMIDA Nygolaimidae Paravulvus hartingii (de Man, 1880) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a).

 Clavicaudoides clavicaudatus (Altherr, 1953) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Solididens vulgaris (Thorne, 1930) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Dorylaimidae Calcaridorylaimus arcticus Gagarin, 1997* Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Calcaridorylaimus promissus Andrássy, 1986 Notes

Alaska (Andrássy 2003c).

 Chrysodorus filiformis (Bastian, 1865) Dorylaimus filiformis Bastian, 1865 Notes

Greenland (Allgén 1954).

 Crocodorylaimus biserovi Gagarin, 1996* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b).

 Crocodorylaimus flavomaculatus (Linstow, 1876) Mesodorylaimus flavomaculatus (Linstow, 1876) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Crocodorylaimus dadayi (Thorne & Swanger, 1936) Laimydorus dadayi (Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986).

 Dorylaimus lineatus Altherr in Altherr & Delamare-Deboutteville, 1972 Notes

Northwest territories, Canada (Mulvey and Anderson 1979).

 Dorylaimus stagnalis Dujardin, 1845 Notes

Greenland (Ditlevsen 1927); Northwest territories, Canada (Mulvey and Anderson 1979); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Laimydorus doryuris (Ditlevsen, 1911) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Laimydorus finalis Thorne, 1975 Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Laimydorus pseudostagnalis (Micoletzly, 1927) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Mesodorylaimus bastiani (Bütschli, 1873) Dorylaimus bastiani Bütschli, 1873; Dorylaimus langii Cobb, 1888 Notes

Svalbard (von Linstow 1900); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Mesodorylaimus japonicus (Cobb in Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Mesodorylaimus litoralis Loof, 1969 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 2001b, Gagarin 2001a).

 Mesodorylaimus pendschikenticus (Tulaganov, 1949) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 2001b, Gagarin 2001a).

 Actinolaimidae Paractinolaimus macrolaimus (de Man, 1884) Actinolaimus macrolaimus (de Man, 1884) Notes

Greenland (Ditlevsen 1927); Alaska (Andrássy 2003c); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Paractinolaimus longidrilus Eveleigh, 1982* Notes

Northwest territories, Canada (Eveleigh 1982).

 Westindicus cinctus (Cobb in Thorne, 1939) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Qudsianematidae Arctidorylaimus arcticus Mulvey & Anderson, 1979* Notes

Northwest territories, Canada (Mulvey and Anderson 1979).

 Allodorylaimus andrassyi (Meyl, 1955) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Allodorylaimus diadematus (Cobb in Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Allodorylaimus holdemani (Andrássy, 1959) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Allodorylaimus lindbergi (Andrássy, 1960) Eudorylaimus lindbergi Andrássy, 1960; Allodorylaimus rarus Gagarin, 1999*; Eudorylaimus curvicaudatus Eliava, 1968 Notes

Lena River estuary, Russia (Gagarin 1999a, Gagarin 1999b, Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 1999a, Gagarin 2001a, Gagarin 2001b); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Allodorylaimus uniformis (Thorne, 1929) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Eudorylaimus acuticauda (de Man, 1880) Dorylaimus acuticauda de Man, 1880; Dorylaimus carteri acuticauda Micoletzky, 1922; Dorylaimus carteri Bastian, 1865 sf. acuticaudata de Man, 1880 (lapsus); Dorylaimus carteri Bastian, 1865 var. brevicaudatus f. typica sf. acuticauda de Man, 1880 Notes

Jan Mayen (Allgén 1953); Greenland (Allgén 1954, Ditlevsen 1927); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Eudorylaimus acutus (Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Eudorylaimus bureshi (Andrássy, 1958) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Eudorylaimus carteri (Bastian, 1865) Dorylaimus carteri Bastian, 1865 Notes

Svalbard (Klekowski and Opalinski 1986, Klekowski and Opalinski 1990, Klekowski and Opalinski 1992, Opalinski and Klekowski 1992); Jan Mayen (Allgén 1953); Greenland (Ditlevsen 1927); Alaska (Andrássy 2003c); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin and Kuzmin 1983, Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Eudorylaimus centrocercus (de Man, 1880) Dorylaimus centrocercus de Man, 1880; Aporcelaimellus centrocercus (de Man, 1880) Notes

Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990).

 Eudorylaimus iners (Bastian, 1865) Dorylaimus gracilis de Man, 1876 Notes

Jan Mayen (Allgén 1953).

 Eudorylaimus leuckarti (Bütschli, 1873) Dorylaimus carteri Bastian, 1865 var. brevicaudatus Micoletzly, 1922 f. typica Micoletzky, 1922 Notes

Greenland (Allgén 1954).

 Eudorylaimus megodon Loof, 1971* Notes

Svalbard (Loof 1971).

 Eudorylaimus subjunctus Loof, 1971* Notes

Svalbard (Loof 1971).

 Eudorylaimus vanrosseni Loof, 1971* Notes

Svalbard (Loof 1971).

 Epidorylaimus agilis (de Man, 1880) Dorylaimus agilis de Man, 1880; Dorylaimus carteri Bastian, 1865 var. agilis de Man, 1880; Eudorylaimus agilis (de Man, 1880); Laimydorus agilis (de Man, 1880) Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Greenland (Allgén 1954); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Epidorylaimus angulosus (Thorne & Swanger, 1936) Eudorylaimus anquilosus (Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Epidorylaimus consobrinus (de Man, 1918) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Kuzmin and Gagarin 1990).

 Epidorylaimus humilis (Thorne & Swanger, 1936) Eudorylaimus humilis (Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Epidorylaimus lugdunensis (de Man, 1880) Dorylaimus carteri Bastian, 1865 sf. lugdunensis de Man, 1880; Dorylaimus carteri lugdunensis Micoletzky, 1922; Eudorylaimus lugdunensis (de Man, 1880); Dorylaimus reisingeri Ditlevsen, 1927* Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Greenland (Ditlevsen 1927); Alaska (Andrássy 2003c); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Epidorylaimus muscorum (Skwarra, 1921) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Epidorylaimus rivalis Gagarin, 1991* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991b, Gagarin 2001b, Gagarin 2001a).

 Microdorylaimus parvus (de Man, 1880) Eudorylaimus parvus (de Man, 1880) Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Crassolabium circulifer (Loof, 1961) Eudorylaimus circulifer Loof, 1961 Notes

Svalbard (Loof 1971).

 Crassolabium ettersbergense (de Man, 1885) Eudorylaimus ettersbergensis (de Man, 1885); Thonus ettersbergensis (de Man, 1885) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Crassolabium laticollis (de Man, 1906) Thonus laticollis (de Man, 1906) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Dorydorella bryophila (de Man, 1880) Dorylaimus carteri Bastian, 1865 sf. briophilus de Man, 1880 Notes

Jan Mayen (Allgén 1953).

 Dorydorella pratensis (de Man, 1880) Dorylaimus carteri Bastian, 1865 sf. pratensis de Man, 1880 Notes

Jan Mayen (Allgén 1953).

 Ecumenicus monohystera (de Man, 1880) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Labronema fimbriatum Thorne, 1939 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Labronema goodeyi Altherr in Altherr & Delamare-Deboutteville, 1972 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 2001b, Gagarin 2001a).

 Labronema loeffleri Andrássy, 1978 Notes

Lena River estuary, Russia (Gagarin 2001b); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a).

 Labronema stechlinense Altherr, 1968 Notes

Alaska (Andrássy 2003c).

 Aporcelaimidae Akrotonus striaticaudatus (Cobb, 1906) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Aporcelaimellus krygeri (Ditlevsen, 1928) Eudorylaimus krygeri (Ditlevsen, 1928) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b, Kuzmin 1972, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 2001b).

 Aporcelaimellus obscurus (Thorne & Swanger, 1936) Eudorylaimus obscurus (Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Aporcelaimellus obtusicaudatus (Bastian, 1865) Dorylaimus obtusicaudatus Bastian, 1865; Dorylaimus obtusicaudatus Bastian, 1865 f. butschlii Micoletzky, 1922; Aporcelaimus obtusicaudatus (Bastian, 1865) Notes

Jan Mayen (Allgén 1953, Allgén 1954); Greenland (Allgén 1954, Ditlevsen 1927); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990).

 Aporcelaimellus papillatus (Bastian, 1865) Aporcelaimus papillatus (Bastian, 1865) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Aporcelaimellus paraobtusicaudatus (Micoletzky, 1922) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Aporcelaimellus tritici (Bastian, 1865) Dorylaimus tritici Bastian, 1865 Notes

Jan Mayen (Allgén 1953); Greenland (Allgén 1954).

 Aporcelaimus eurydoris (Ditlevsen, 1911) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Aporcelaimus polaris Andrássy, 2003* Notes

Alaska (Andrássy 2003c).

 Aporcelaimus superbus (de Man, 1880) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Metaporcelaimus labiatus (de Man, 1880) Dorylaimus labiatus de Man, 1880 Notes

Jan Mayen (Allgén 1953).

 Crateronematidae Chrysonema maksymovi (Altherr, 1963) Eudorylaimus maksymovi (Altherr, 1963) Notes

Svalbard (Klekowski and Opalinski 1986, Klekowski and Opalinski 1990, Loof 1971).

 Nordiidae Enchodelus analatus (Ditlevsen, 1927) Dorylaimus (Doryllium) analatus Ditlevsen, 1927* Notes

Svalbard (Loof 1971); Greenland (Ditlevsen 1927).

 Enchodelus groenlandicus (Ditlevsen, 1927) Dorylaimus (Doryllium) groenlandicus Ditlevsen 1927* Notes

Greenland (Ditlevsen 1927); Taymyr and Severnaya Zemlya, Russia (Elshishka et al. 2012).

 Enchodelus cf. macrodoroides (Steiner, 1914) Notes

Svalbard (Loof 1971).

 Enchodelus macrodorus (de Man, 1880) Dorylaimus macrodorus de Man, 1880 Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b, Steiner 1916a).

 Enchodelus makarovae Elshishka et al., 2012* Notes

Taymyr and Severnaya Zemlya, Russia (Elshishka et al. 2012).

 Enchodelus parateres Baqri & Jairajpuri, 1974 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Enchodelus parvus Loof, 1971* Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Enchodelus teres Thorne, 1939 Notes

Taymyr and Severnaya Zemlya, (Kuzmin and Gagarin 1990).

 Heterodorus arquatus (Thorne, 1939) Enchodelus arquatus Thorne, 1939 Notes

Nunavut, Canada (Elshishka et al. 2013); Taymyr and Severnaya Zemlya, Russia (Elshishka et al. 2013, Kuzmin and Gagarin 1990).

 Heterodorus conicaudatus (Ditlevsen, 1927) Dorylaimus conicaudatus Ditlevsen, 1927*; Enchodelus conicaudatus (Ditlevsen, 1927) Notes

Svalbard (Loof 1971); Greenland (Ditlevsen 1927).

 Heterodorus magnificus Altherr, 1952 Notes

Taymyr and Severnaya Zemlya, Russia (Elshishka et al. 2013).

 Pungentus longidens (Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Rhyssocolpus alleni (Brzeski, 1962) Eudorylaimus alleni Brzeski, 1962* Notes

Svalbard (Brzeski 1962, Loof 1971).

 Rhyssocolpus aquilonius Andrássy, 2003* Notes

Alaska (Andrássy 2003c).

 Rhyssocolpus arcticus Ebsary, 1984* Notes

Nunavut, Canada (Ebsary 1984).

 Longidorella magna Loof, 1971* Notes

Svalbard (Loof 1971); Alaska (Andrássy 2003c).

 Longidorella parva Thorne, 1939 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Longidorella penetrans (Thorne & Swanger, 1936) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Leptonchidae Xiphinemella eversum Heyns, 1969 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Tylencholaimidae Tylencholaimus crassus Loof & Jairajpuri, 1968 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Tylencholaimus mirabilis (Bütschli, 1873) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Tylencholaimus proximus Thorne, 1939 Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Tylencholaimus stecki Steiner, 1914 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Tylencholaimus teres Thorne, 1939 Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990).

 Mydonomidae Dorylaimoides arcuicaudatus Baqri & Jairajpuri, 1969 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Tylencholaimellidae Tylencholaimellus coronatus Thorne, 1939 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 MONONCHIDA Mononchidae Mononchus absconditus (Tsalolikhin, 1974) Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 2001b, Gagarin 2001a); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Mononchus angarensis Gagarin, 1984 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b).

 Mononchus aquaticus Coetzee, 1968 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b).

 Mononchus maduei Schneider, 1925 Notes

Northwest territories, Canada (Mulvey 1978).

 Mononchus niddensis Skwarra, 1921 Notes

Nunavut, Canada (Mulvey 1967b, Mulvey 1978).

 Mononchus superbus Mulvey, 1978* Notes

Northwest territories, Canada (Mulvey 1978).

 Mononchus tajmiris Gagarin, 1991* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1991b, Gagarin 1993, Gagarin 2001a, Gagarin 2001b).

 Mononchus truncatus Bastian, 1865 Notes

Northwest territories, Canada (Mulvey 1978); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Paramononchus arcticus Mulvey, 1978* Notes

Northwest territories, Canada (Mulvey 1978).

 Prionchulus longus (Thorne, 1929) Notes

Nunavut, Canada (Mulvey 1967a, Mulvey 1978); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Prionchulus major Gagarin, 2001* Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 2001c, Gagarin 2001b).

 Prionchulus muscorum (Dujardin, 1845) Notes

Svalbard (Klekowski and Opalinski 1986, Klekowski and Opalinski 1990); Nunavut, Canada (Mulvey 1967a); Alaska (Andrássy 2003a); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Prionchulus punctatus Cobb, 1917 Notes

Nunavut, Canada (Mulvey 1967a).

 Prionchulus sedatus Gagarin, 2000* Notes

Lena River estuary, Russia (Gagarin 2000, Gagarin 2001b).

 Prionchulus spectabilis (Ditlevsen, 1911) Mononchus spectabilis Ditlevsen, 1911 Notes

Greenland (Ditlevsen 1927).

 Clarkus macropapillatus (Mulvey, 1967) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Clarkus papillatus (Bastina, 1865) Mononchus papillatus Bastian, 1865 Notes

Jan Mayen (Allgén 1953); Greenland (Allgén 1954, Ditlevsen 1927); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Coomansus fletcherensis Mulvey, 1978* Notes

Nunavut, Canada (Mulvey 1978).

 Coomansus gerlachei (de Man, 1904) Notes

Northwest territories, Canada (Mulvey 1978).

 Coomansus parvus (de Man, 1880) Notes

Alaska (Andrássy 2003a).

 Parkellus zschokkei (Menzel, 1913) Iotonchus zschokkei (Menzel, 1913) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Mylonchulidae Mylonchulus brachyuris (Bütschli, 1873) Mononchus brachyuris Bütschli, 1873 Notes

Greenland (Ditlevsen 1927); Alaska (Andrássy 2003a); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Mylonchulus gigas Gagarin, 1993 Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Mylonchulus incurvus Cobb, 1917 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b).

 Mylonchulus lacustris (Cobb in Cobb, 1915) Notes

Northwest territories, Canada (Mulvey 1978).

 Anatonchidae Miconchus exilis (Cobb, 1917) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Iotonchidae Jensenonchus amplus Andrássy, 2003* Notes

Alaska (Andrássy 2003a).

 CHROMADORIDA Chromadoridae Chromadorita arctica Gagarin, 1999* Chromadorida arctica Gagarin, 1999 (lapsus) Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1999a, Gagarin 2001b).

 Chromadorita leuckarti (de Man, 1876) Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Punctodora ratzemburgensis (Linstow, 1876) Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Achromadoridae Achromadora micoletzkyi (Stefanski, 1915) Cyatholaimus micoletzkyi Steiner, 1916*; Achromadora steineri Mulvey, 1969 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Achromadora ruricola (de Man, 1880) Notes

Nunavut, Canada (Mulvey 1969c); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Achromadora cf. semiarmata Altherr, 1952 Notes

Svalbard (Loof 1971).

 Achromadora tenax (de Man, 1876) Notes

Svalbard (Janiec 1997, Loof 1971).

 Achromadora terricola (de Man, 1880) Cyatholaimus ornatus Steiner, 1916* Notes

Greenland (Ditlevsen 1927); Nunavut, Canada (Mulvey 1969c); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b, Steiner 1916a).

 Ethmolaimidae Ethmolaimus pratensis de Man, 1880 Ethmolaimus arcticus Steiner, 1916* Notes

Nunavut, Canada (Mulvey 1969c); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Kuzmin 1972, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b, Steiner 1916a).

 DESMODORIDA Microlaimidae Microlaimus globiceps de Man, 1880 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Prodesmodora arctica (Mulvey, 1969) Microlaimus arcticus Mulvey 1969* Notes

Nunavut, Canada (Mulvey 1969c); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Prodesmodora circulata (Micoletzky, 1913) Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 MONHYSTERIDA Xyalidae Hofmaenneria hazanensis Mulvey, 1969* Notes

Nunavut, Canada (Mulvey 1969c).

 Daptonema dubium (Bütschli, 1873) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin and Kuzmin 1983, Kuzmin and Gagarin 1990).

 Daptonema fortis Gagarin, 1993* Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 2001b, Gagarin 2001a).

 Daptonema sibiricum Gagarin, 2000* Notes

Lena River estuary, Russia (Gagarin 2000, Gagarin 2001b).

 Daptonema tenuispiculum (Ditlevsen, 1918) Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1999a, Gagarin 2001b).

 Theristus agilis (de Man, 1880) Monhystera agilis de Man, 1880 Notes

Jan Mayen (Allgén 1953, Tchesunov and Pletnikova 1986); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990).

 Theristus flevensis Schuurmans Stekhoven, 1935 Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997b, Gagarin 2001a).

 Monhysteridae Eumonhystera barbata Andrássy, 1981 Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997b, Gagarin 2001a).

 Eumonhystera dispar (Bastian, 1865) Monhystera dispar Bastian, 1865 Notes

Svalbard (Loof 1971); Nunavut, Canada (Mulvey 1969c); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997b, Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Eumonhystera filiformis (Bastian, 1865) Monohystera filiformis Bastian, 1865 (lapsus); Monhystera filiformis Butschlii, 1873 (lapsus) Notes

Svalbard (Janiec 1997, Loof 1971); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b, Steiner 1916a).

 Eumonhystera kuzmini Gagarin, 1997* Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997b, Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Eumonhystera maxima Gagarin, 1996 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b).

 Eumonhystera papuana (Daday, 1899) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b).

 Eumonhystera pseudobulbosa (Daday, 1896) Monohystera pseudobulbosa Daday, 1896 (lapsus) Notes

Greenland (Ditlevsen 1927).

 Eumonhystera similis (Bütschli, 1873) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b, Kuzmin and Gagarin 1990).

 Eumonhystera simplex (de Man, 1880) Monhystera simplex de Man, 1880 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Eumonhystera tuporis Gagarin, 1991 Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1999a, Gagarin 2001b).

 Eumonhystera vulgaris (de Man, 1880) Monhystera vulgaris de Man, 1880; Monohystera vulgaris de Man, 1880 (lapsus) Notes

Svalbard (Janiec 1997, Loof 1971); Jan Mayen (Allgén 1953); Nunavut, Canada (Mulvey 1969c); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b, Steiner 1916a).

 Geomonhystera aenariensis (Meyl, 1953) Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Geomonhystera paravillosa (Meyl, 1954) Monhystera paravillosa Meyl, 1954 Notes

Nunavut, Canada (Mulvey 1969c).

 Geomonhystera villosa (Butschli, 1873) Monohystera villosa Butschli, 1873 (lapsus) Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Halomonhystera disjuncta (Bastian, 1865) Monhystera disjuncta Bastian, 1865 Notes

Jan Mayen (Allgén 1953); Novaya Zemlya and Vaigach island, Russia (Gagarin 1999a, Gagarin 2001b).

 Monhystera amabilis Gagarin, 1997* Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997b, Gagarin 1997a, Gagarin 2001c, Gagarin 2001b).

 Monhystera stagnalis Bastian, 1865 Notes

Svalbard (Janiec 1997, Loof 1971); Taymyr and Severnaya Zemlya, Russia (Gagarin and Kuzmin 1983, Kuzmin and Gagarin 1990).

 Monhystrella macrura (de Man, 1880) Monhystera macrura de Man, 1880 Notes

Jan Mayen (Allgén 1953).

 Tridentula bidenticulata (Gagarin, 1997) Eumonhystera bidenticulata Gagarin, 1997*; Tridentulus diplodenticulata Gagarin, 1997 (lapsus); Eumonhystera diplodenticulata Gagarin, 1997 (lapsus) Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997b, Gagarin 1997a, Gagarin 2001b).

 Tridentula obscura (Gagarin, 2000) Tridentulus obscurus Gagarin, 2000 Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 2001b).

 ARAEOLAIMIDA Diplopeltidae Cylindrolaimus baradlanus Andrássy, 1959 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Cylindrolaimus communis de Man, 1880 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Cylindrolaimus melancholicus de Man, 1880 Notes

Svalbard (Loof 1971); Nunavut, Canada (Mulvey 1969c); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 PLECTIDA Camacolaimidae Deontolaimus papillatus de Man, 1880 Notes

Greenland (Ditlevsen 1927).

 Chronogastridae Chronogaster typica (de Man, 1921) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Plectidae Anaplectus grandepapillatus (Ditlevsen, 1928) Anaplectus submersus (Hirschmann, 1952) Notes

Svalbard (Janiec 1997); Alaska (Andrássy 2003b); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990).

 Anaplectus granulosus (Bastian, 1865) Plectus granulosus Bastian, 1865 Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Greenland (Ditlevsen 1927); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b, Steiner 1916a).

 Anaplectus porosus Allen and Noffsinger, 1968 Notes

Svalbard (Klekowski and Opalinski 1986, Klekowski and Opalinski 1990, Klekowski and Opalinski 1992, Loof 1971).

 Arctiplectus alaskanus Andrássy, 2003* Notes

Alaska (Andrássy 2003b).

 Perioplectus labiosus (Sanwal, 1968) Periplectus labiosus Sanwal, 1968* Notes

Nunavut, Canada (Sanwal 1968, Zell 1993).

 Perioplectus secundus Andrássy, 2003* Notes

Alaska (Andrássy 2003b).

 Plectus acuminatus Bastian, 1865 Notes

Svalbard (Loof 1971); Alaska (Andrássy 2003b); Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Plectus amorphotelus Ebsary, 1985 Notes

Svalbard (Zell 1993).

 Plectus aquatilis Andrássy, 1984 Notes

Svalbard (Janiec 1997).

 Plectus cf. armatus Butschli, 1873 Notes

Svalbard (Loof 1971).

 Plectus assimilis Bütschli, 1873 Ceratoplectus assimilis (Bütschli, 1873) Notes

Svalbard (Loof 1971); Greenland (Zell 1993); Alaska (Andrássy 2003b); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Plectus cancellatus Zullini, 1978 Plectus thornei sensu Zell, 1993 Notes

Nunavut, Canada (Ebsary 1985, Zell 1993).

 Plectus cirratus Bastian, 1865 Notes

Svalbard (Klekowski and Opalinski 1986, Klekowski and Opalinski 1990, Klekowski and Opalinski 1992, Menzel 1920); Jan Mayen (Allgén 1953); Greenland (Ditlevsen 1927); Alaska (Andrássy 2003b); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Plectus communis Bütschli, 1873 Notes

Svalbard (Menzel 1920, Zell 1993); Iceland (Zell 1993); Greenland (Allgén 1954).

 Plectus cornus Maggenti, 1961 Ceratoplectus cornus (Maggenti, 1961) Notes

Svalbard (Loof 1971, Zell 1993); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1976, Kuzmin and Gagarin 1990).

 Plectus elongatus Maggenti, 1961 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Plectus geophilus de Man, 1880 Plectus minor Novikova & Gagarin, 1971 Notes

Svalbard (Loof 1971, Zell 1993); Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Plectus inquirendus Andrássy, 1958 Notes

Svalbard (Loof 1971, Zell 1993); Alaska (Andrássy 2003b); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Plectus intermedius Cobb, 1893 Notes

Nunavut, Canada (Zell 1993).

 Plectus longicaudatus Butschli, 1873 Notes

Svalbard (Loof 1971, Zell 1993); Jan Mayen (Allgén 1953); Iceland (Zell 1993); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Plectus magadani Kuzmin, 1979 Notes

Svalbard (Zell 1993).

 Plectus murrayi Yeates, 1970 Notes

Svalbard (Zell 1993).

 Plectus cf. opisthocirculus Andrássy, 1952 Notes

Svalbard (Janiec 1997).

 Plectus palustris de Man, 1880 Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Plectus parietinus Bastian, 1865 Plectus cirratus Bastian, 1865 f. parietinus Bastian, 1865 Notes

Svalbard (Janiec 1997, Klekowski and Opalinski 1986, Klekowski and Opalinski 1990, Klekowski and Opalinski 1992, Loof 1971, Opalinski and Klekowski 1992); Jan Mayen (Allgén 1953); Iceland (Zell 1993); Alaska (Andrássy 2003b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1996, Gagarin 2001b, Gagarin 2001a, Gagarin and Kuzmin 1983, Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b, Steiner 1916a).

 Plectus parvus Bastian, 1865 Notes

Svalbard (Loof 1971, Zell 1993); Iceland (Zell 1993); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Plectus rhizophilus de Man, 1880 Plectus cirratus Bastian, 1865 var. rhizophilus de Man, 1880 Notes

Svalbard (Loof 1971, Zell 1993); Jan Mayen (Allgén 1953, Steiner 1916a); Greenland (Allgén 1954); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin and Kuzmin 1983, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b, Steiner 1916a).

 Plectus rotundilabiatus Zell, 1993 Notes

Iceland (Zell 1993).

 Plectus tenuis Bastian, 1865 Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 1999a, Gagarin 2001b).

 Plectus varians Maggenti, 1961 Notes

Iceland (Zell 1993); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1976).

 Plectus velox Bastian, 1865 Notes

Svalbard (Zell 1993).

 Tylocephalus auriculatus (Bütschli, 1873) Plectus auriculatus Bütschli, 1873 Notes

Jan Mayen (Steiner 1916a); Nunavut, Canada (Anderson 1966); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Ereptonema arcticum Loof, 1971* Notes

Svalbard (Boström 1987a, Holovachov et al. 2003, Loof 1971).

 Ereptonema fimbriatum Anderson, 1966* Notes

Nunavut, Canada (Anderson 1966).

 Wilsonema otophorum (de Man, 1880) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Metateratocephalidae Metateratocephalus crassidens (de Man, 1880) Teratocephalus crassidens de Man, 1880; Euteratocephalus crassidens (de Man, 1880) Notes

Svalbard (Boström 1989, Loof 1971); Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b, Steiner 1916a).

 Euteratocephalus palustris (de Man, 1880) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Aulolaimidae Aulolaimus oxycephalus de Man, 1880 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 RHABDITIDA Teratocephalidae Teratocephalus costatus Andrássy, 1958 Teratocephalus decarinus Anderson, 1969* Notes

Svalbard (Boström 1989, Loof 1971); Nunavut, Canada (Anderson 1969); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Teratocephalus dadayi Andrássy, 1968 Teratocephalus subvexus Anderson, 1969* Notes

Nunavut, Canada (Anderson 1969).

 Teratocephalus lirellus Anderson, 1969* Notes

Svalbard (Boström 1989, Loof 1971); Nunavut, Canada (Anderson 1969); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Teratocephalus tenuis Andrássy, 1958 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Teratocephalus terrestris (Bütschli, 1873) Notes

Svalbard (Boström 1989); Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Alloionematidae Rhabditophanes schneideri (Bütschli, 1873) Cheilobus quadrilabiatus Cobb, 1924 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Panagrolaimidae Panagrolaimus papillosus Loof, 1971* Notes

Svalbard (Loof 1971).

 Panagrolaimus rigidus (Schneider, 1866) Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1976, Kuzmin and Gagarin 1990).

 Cephalobidae Cephalobus filiformis de Man, 1876 Cephalobus filiformis species inquirenda Notes

Jan Mayen (Allgén 1953).

 Cephalobus persegnis Bastian, 1865 Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Acrobeloides buetschlii (de Man, 1884) Cephalobus buetschlii de Man, 1884 Notes

Svalbard (Menzel 1920); Jan Mayen (Steiner 1916a); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Acrobeloides enoploides Loof, 1971* Notes

Svalbard (Loof 1971).

 Acrobeloides nanus (de Man, 1880) Cephalobus nanus de Man, 1880 Notes

Svalbard (Loof 1971).

 Acrobeloides tricornis (Thorne, 1925) Notes

Svalbard (Boström 1987a, Loof 1971).

 Eucephalobus arcticus Loof, 1971* Notes

Svalbard (Boström 1987a, Loof 1971).

 Eucephalobus mucronatus (Kozlovska & Roguska-Wasilewska, 1963) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Eucephalobus oxyuroides (de Man, 1876) Cephalobus oxyuroides de Man, 1876 Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953).

 Eucephalobus paracornutus De Coninck, 1943 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Eucephalobus striatus (Bastian, 1865) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Pseudacrobeles elongatus (de Man, 1880) Cephalobus elongatus de Man, 1880; Heterocephalobus elongatus (de Man, 1880) Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b, Kuzmin 1972, Kuzmin and Gagarin 1990).

 Chiloplacus lentus (Maupas, 1900) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Chiloplacus quintastriatus Sumenkova in Sumenkova & Razjivin, 1968 Chiloplacus saccatus Loof, 1971* Notes

Svalbard (Boström 1987a, Loof 1971).

 Chiloplacus symmetricus (Thorne, 1925) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Chiloplacus trilineatus Steiner, 1940 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Stegelletina insubrica (Steiner, 1914) Cervidellus insubricus (Steiner, 1914); Ypsylonellus insubricus (Steiner, 1914) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Stegelletina similis (Thorne, 1925) Stegelleta mucronata Loof, 1971* Notes

Svalbard (Loof 1971).

 Cervidellus vexilliger (de Man, 1880) Cervidellus serratus (Thorne, 1925); Ypsylonellus vexilliger (de Man, 1880) Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Cervidellus spitzbergensis Boström, 1987* Notes

Svalbard (Boström 1987b).

 Acrobeles ciliatus von Linstow, 1877 Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Aphelenchoididae Aphelenchoides arcticus Sanwal, 1965* Notes

Svalbard (Loof 1971); Nunavut, Canada (Sanwal 1965); Lena River estuary, Russia (Gagarin 2001b).

 Aphelenchoides blastophthorus Franklin, 1952 Notes

Lena River estuary, Russia (Gagarin 2001b).

 Aphelenchoides echinocaudatus Haque, 1968 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Aphelenchoides goeldii (Steiner, 1914) Aphelenchus goeldii Steiner, 1914 Notes

Jan Mayen (Steiner 1916a).

 Aphelenchoides parasaprophilus Sanwal, 1965* Notes

Nunavut, Canada (Sanwal 1965); Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b).

 Aphelenchoides parietinus (Bastian, 1865) Aphelenchus modestus de Man, 1876 Notes

Jan Mayen (Steiner 1916a); Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Aphelenchoides robustus Gagarin, 1997* Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997c, Gagarin 1997a, Gagarin 2001b).

 Aphelenchoides saprophilus Franklin, 1957 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Aphelenchoides subtenuis (Cobb, 1926) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Tylenchidae Aglenchus agricola (de Man, 1884) Notes

Greenland (Brzeski 1999); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Coslenchus costatus (de Man, 1921) Tylenchus costatus de Man, 1921 Notes

Svalbard (Loof 1971); Greenland (Brzeski 1999); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Basiria dolichura Loof, 1971* Notes

Svalbard (Loof 1971).

 Lelenchus leptosoma (de Man, 1880) Tylenchus leptosoma de Man, 1880; Anguillulina leptosoma (de Man, 1880); Filenchus leptosoma (de Man, 1880) Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Filenchus cylindricaudus (Wu, 1969) Tylenchus cylindricaudus Wu, 1969* Notes

Nunavut, Canada (Wu 1969b).

 Filenchus cylindricus (Thorne & Malek, 1968) Notes

Lena River estuary, Russia (Gagarin 2001b).

 Filenchus discrepans (Andrássy, 1954) Lelenchus discrepans Andrássy, 1954; Ottolenchus discrepans (Andrássy, 1954) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Filenchus ditissimus (Brzeski, 1963) Tylenchus ditissimus Brzeski, 1963 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin and Kuzmin 1983, Kuzmin and Gagarin 1990).

 Filenchus facultativus (Szczygiel, 1970) Notes

Greenland (Brzeski 1999).

 Filenchus filiformis (Bütschli, 1873) Filenchus filiformis species inquirenda; Tylenchus filiformis Bütschli, 1873; Tylenchus (Filenchus) filiformis Bütschli, 1873 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Filenchus hamatus (Thorne & Malek, 1968) Notes

Greenland (Brzeski 1999).

 Filenchus hazanensis (Wu, 1969) Tylenchus hazenensis Wu, 1969*; Dactylotylenchus filiformis Wu, 1969* Notes

Nunavut, Canada (Wu 1969b, Wu 1969a).

 Filenchus minutus (Cobb, 1893) Filenchus minutus species inquirenda Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Filenchus misellus (Andrássy, 1958) Notes

Greenland (Brzeski 1999).

 Filenchus orbus Andrássy, 1954 Tylenchus aquilonius Wu, 1969* Notes

Greenland (Brzeski 1999); Nunavut, Canada (Wu 1969b).

 Filenchus quartus (Szczygiel, 1969) Notes

Greenland (Brzeski 1999).

 Filenchus thornei (Andrássy, 1954) Tylenchus thornei Andrássy, 1954 Notes

Svalbard (Loof 1971); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Malenchus acarayensis Andrássy, 1968 Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Malenchus bryophilus (Steiner, 1914) Tylenchus bryophilus Steiner, 1914 Notes

Svalbard (Loof 1971); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b).

 Malenchus exiguus (Massey, 1969) Ottolenchus sulcis Wu, 1970* Notes

Nunavut, Canada (Wu 1970).

 Tylenchus davainei Bastian, 1865 Anguillulina davainei (Bastian, 1865); Filenchus davainei (Bastian, 1865) Notes

Svalbard (Loof 1971); Jan Mayen (Allgén 1953); Greenland (Brzeski 1999, Ditlevsen 1927); Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin and Kuzmin 1983, Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2001b, Steiner 1916a).

 Tylenchus neodavainei Wu, 1969* Notes

Nunavut, Canada (Wu 1969b).

 Tylenchus stylolineatus Wu, 1969* Notes

Nunavut, Canada (Wu 1969b).

 Dolichodoridae Meiodorus hyalacus (Anderson & Ebsary, 1982) Mulveyotus hyalacus Anderson & Ebsary, 1982* Notes

Nunavut, Canada (Anderson and Ebsary 1982).

 Geocenamus arcticus (Mulvey, 1969) Tylenchorhynchus arcticus Mulvey, 1969* Notes

Svalbard (Loof 1971); Nunavut, Canada (Mulvey 1969a); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Geocenamus brevidens (Allen, 1955) Merlinius brevidens (Allen, 1955) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Geocenamus circellus (Anderson & Ebsary, 1982) Merlinius circellus Anderson & Ebsary, 1982* Notes

Nunavut, Canada (Anderson and Ebsary 1982).

 Geocenamus joctus (Thorne, 1949) Tetylenchus joctus Thorne, 1949 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Geocenamus loofi (Siddiqi, 1979) Tetylenchus joctus sensu Loof, 1971; Merlinius loofi Siddiqi, 1979 Notes

Svalbard (Loof 1971, Siddiqi 1979).

 Geocenamus microdorus (Geraert, 1966) Tylenchorhynchus microdorus Geraert, 1966 Notes

Svalbard (Loof 1971).

 Geocenamus tetylus (Anderson & Ebsary, 1982) Merlinius tetylus Anderson & Ebsary, 1982* Notes

Nunavut, Canada (Anderson and Ebsary 1982).

 Nagellus borealis Powers, Baldwin & Bell, 1983* Notes

Alaska (Powers et al. 1983).

 Nagellus grandis (Allen, 1955) Tylenchorhynchus grandis Allen, 1955 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Nagelus leptus (Allen, 1955) Tylenchorhynchus leptus Allen, 1955 Notes

Svalbard (Loof 1971); Greenland (Brzeski 1999); Nunavut, Canada (Mulvey 1969a); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Nagellus lineatus (Allen, 1955) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Nagellus obscurus (Allen, 1955) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Nagellus parobscurus (Mulvey, 1969) Tylenchorhynchus parobscurus Mulvey, 1969* Notes

Nunavut, Canada (Mulvey 1969a).

 Tylenchorhynchus dubius (Bütschli, 1873) Bitylenchus dubius (Bütschli, 1873) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin 1976, Kuzmin and Gagarin 1990).

 Tylenchorhynchus nanus Allen, 1955 Geocenamus nanus (Allen, 1955); Merlinius nanus (Allen, 1955) Notes

Greenland (Brzeski 1999); Taymyr and Severnaya Zemlya, Russia (Kuzmin 1986, Kuzmin and Gagarin 1990).

 Tylenchorhynchus cf. nothus Allen, 1955 Notes

Svalbard (Loof 1971).

 Tylenchorhynchus parvus Allen, 1955 Bitylenchus parvus (Allen, 1955) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Hoplolaimidae Rotylenchus brevicaudatus (Hopper, 1959) Pararotylenchus brevicaudatus (Hopper, 1959) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Helicotylenchus platyurus Perry in Perry, Darling & Thorne, 1959 Notes

Greenland (Brzeski 1999).

 Helicotylenchus spitsbergensis Loof, 1971* Notes

Svalbard (Loof 1971); Greenland (Brzeski 1999).

 Helicotylenchus varicaudatus Yuen, 1964 Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Pratylenchidae Pratylenchoides crenicauda Winslow, 1958 Notes

Svalbard (Loof 1971).

 Pratylenchoides magnicauda (Thorne, 1935) Tylenchorhynchus magnicauda (Thorne, 1935) Notes

Svalbard (Loof 1971).

 Pratylenchus pratensis (de Man, 1880) Anguillulina pratense (de Man, 1880) Notes

Jan Mayen (Allgén 1953).

 Hirschmanniella gracilis (de Man, 1880) Tylenchus gracilis Cobb (lapsus) Notes

Svalbard (von Linstow 1900).

 Hirschmanniella trimucronata Gagarin, 2000* Notes

Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a, Gagarin 2000, Gagarin 2001b).

 Criconematidae Criconemoides annulatus Cobb in Taylor, 1936 Criconemoides hemispaericaudatus Wu, 1965* Notes

Svalbard (Loof 1971); Nunavut, Canada (Wu 1965).

 Tylenchulidae Paratylenchus macrophallus (de Man, 1880) Paratylenchus macrophallus species inquirenda; Anguillulina macrophallum (de Man, 1880) Notes

Jan Mayen (Allgén 1953).

 Paratylenchus cf. projectus Jenkins, 1956 Notes

Greenland (Brzeski 1999).

 Anquinidae Anguina agrostis (Steinbuch, 1799) Notes

Nunavut, Canada (Mulvey 1963).

 Subanguina askenasyi (Bütschli, 1873) Ditylenchus askenasyi (Bütschli, 1873) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b).

 Ditylenchus acutatus Brzeski, 1991 Notes

Greenland (Brzeski 1999).

 Ditylenchus dryadis Anderson & Mulvey, 1980* Notes

Nunavut, Canada (Anderson and Mulvey 1980).

 Ditylenchus equalis Heyns, 1964 Notes

Greenland (Brzeski 1999).

 Ditylenchus intermedius (de Man, 1880) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Nothotylenchus acris Thorne, 1941 Notes

Nunavut, Canada (Mulvey 1969b); Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Nothotylenchus acutus Khan, 1965 Notes

Nunavut, Canada (Mulvey 1969b).

 Nothotylenchus attenuatus Mulvey, 1969* Notes

Nunavut, Canada (Mulvey 1969b).

 Nothotylenchus cf. danubialis Andrássy, 1960 Notes

Nunavut, Canada (Mulvey 1969b).

 Neotylenchidae Deladenus durus (Cobb, 1922) Notes

Nunavut, Canada (Das 1964).

 Hadrodenus megacondylus Mulvey, 1969* Notes

Nunavut, Canada (Mulvey 1969b).

 Hexatylus mulveyi Das, 1964* Notes

Nunavut, Canada (Das 1964).

 Prothallonema mucronatum (Thorne & Malek, 1968) Stictylus mucronatus Thorne & Malek, 1968 Notes

Greenland (Brzeski 1999); Nunavut, Canada (Mulvey 1969b).

 Bunonematidae Bunonema hessi Steiner, 1914 Notes

Novaya Zemlya and Vaigach island, Russia (Steiner 1916b).

 Bunonema reticulatum Richters, 1905 Notes

Svalbard (Loof 1971).

 Diplogastridae Diplogaster rivalis (Leydig, 1854) Notes

Lena River estuary, Russia (Gagarin 2001b).

 Goffartia variabilis (Micoletzky, 1922) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 2001b, Gagarin 2001a, Gagarin 2008).

 Fictor fictor (Bastian, 1865) Notes

Taymyr and Severnaya Zemlya, Russia (Kuzmin and Gagarin 1990).

 Fictor shoshini Gagarin, 1995* Notes

Lena River estuary, Russia (Gagarin 1995, Gagarin 2001b); Novaya Zemlya and Vaigach island, Russia (Gagarin 1997a).

 Koerneria angarensis (Gagarin, 1983) Fictor angarensis (Gagarin, 1983) Notes

Lena River estuary, Russia (Gagarin 2001b); Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 2001b, Gagarin 2001a).

 Koerneria ruricola (Gagarin, 1983) Fictor ruricola (Gagarin, 1983) Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 2001b).

 Mononchoides intermedius Gagarin, 1993 Notes

Taymyr and Severnaya Zemlya, Russia (Gagarin 1996, Gagarin 2001b).

 Rhabditidae Choriorhabditis gracilicauda (de Man, 1876) Rhabditis gracilicauda de Man, 1876 Notes

Jan Mayen (Allgén 1953).

 Choriorhabditis producta (Schneider, 1866) Anguillulina intermedia (de Man, 1880) Notes

Jan Mayen (Allgén 1953).

 Rhabditis terricola Dujardin, 1844 Rhabditis aspera Bütschli, 1873 Notes

Jan Mayen (Allgén 1953).

 Aphelenchus nivalis Aurivillius, 1883* Notes

Inquirenda et incertae sedis. Svalbard (Aurivillius 1883).

 Macfadyenia filicaudata Allgén, 1953* Notes

Inquirenda et incertae sedis. Jan Mayen (Allgén 1953).

 Parachromagasteriella arctica Allgén, 1953* Notes

Inquirenda et incertae sedis. Jan Mayen (Allgén 1953).

 Plectus cirratusf.parietinus sf. latilaimus Allgén, 1953* Notes

Inquirenda et incertae sedis. Jan Mayen (Allgén 1953).

 Prodomorganus tajmiris Gagarin, 1993* Notes

Inquirenda et incertae sedis. Taymyr and Severnaya Zemlya, Russia (Gagarin 1990, Gagarin 1993, Gagarin 2001b).

 Teratocephalus micrurus Rahm, 1924 Notes

Inquirenda et incertae sedis. Taymyr and Severnaya Zemlya, Russia (Kuzmin 1972, Kuzmin and Gagarin 1990).

 Tylenchus macrospiculatus Gagarin, 1997 Notes

Nomen nudum. Novaya Zemlia and Vaigach island, Russia (Gagarin 1997a).

 Discussion

Despite of more than 130 years of research, our knowledge on the diversity and distribution of Arctic nematodes is still very incomplete and limited to only few relatively small areas of the region. The first studies of Arctic nematodes date back to 1883, when the first nematode was described from Spitzbergen (Aurivillius 1883). The taxonomic position of the species Aphelenchus nivalis Aurivillius, 1883 remains uncertain and its type material can not be located for re-study. After Aurivillius, the nematode fauna of Svalbard archipelago was studied by von Linstow (1900), Menzel (1920), Boström (1987a), Boström (1987b), Boström (1989), Zell (1993) and to a lesser extent by Brzeski (1962), Siddiqi (1979), Janiec (1997), Klekowski and Opalinski (1986), Klekowski and Opalinski (1990), Klekowski and Opalinski (1992), Opalinski and Klekowski (1992), Holovachov et al. (2003) adding up to the list of 101 species. Most of the sampling on Svalbard was concentrated around the west part of the Spitsbergen island and a large part of the archipelago remains unstudied. From other territories belonging to the Kingdom of Norway, 43 species of nematodes were found on the small island of Jan Mayen (Allgén 1953, Steiner 1916a), while no information is available for the northernmost part of the Finnmark.

Most of the nematological research in the Iceland was focused in southern and western part of the island (De Coninck 1940, De Coninck 1943, De Coninck 1944), which is considered as Subarctic in the current publication. Sampling sites from the northern part of the country were also outside the boundary considered in the present paper (Kreis 1963, Starmühlner 1969). A small part of the Arctic of the Iceland has not been studied to any considerable extent – only six species from the genus Plectus are listed from the areas near Húsavík and Tjörnes by Zell (1993). Terrestrial nematode fauna of Greenland, on the other hand, received considerable attention from Ditlevsen (1927), Allgén (1954), Zell (1993), Brzeski (1999), who altogether reported 43 species of nematodes collected in several distant localities along the eastern (Little Pendulum Island and Angmagssalik) and western (Disko island, Kangerlussuaq, Kapisillit, Sukkertoppen and Upernavik) sides of the island.

Nematode fauna of the Arctic regions of Canada is studied comparatively better. The research was focused in several localities in the Nunavut province including Bathhurst, Ellef Ringnes, Ellesmere and Somerset islands (Anderson 1966, Anderson 1969, Anderson and Ebsary 1982, Anderson and Mulvey 1980, Das 1964, Ebsary 1984, Ebsary 1985, Elshishka et al. 2013, Mulvey 1963, Mulvey 1967b, Mulvey 1967a, Mulvey 1969c, Mulvey 1969a, Mulvey 1969b, Mulvey 1978, Sanwal 1965, Sanwal 1968, Wu 1965, Wu 1969b, Wu 1969a, Wu 1970, Zell 1993), and in the small area of the Mackenzie River delta and Tuktoyaktok in the Northwest territories (Ebsary 1982, Eveleigh 1982, Mulvey 1978, Mulvey and Anderson 1979). Respectively, 55 and 13 species were described from these regions between 1963 and 1993. Much less is known about the nematodes from the Arctic regions of Alaska – only 22 species were recorded there by Andrássy (2003a), Andrássy (2003b), Andrássy (2003c), Bernard (1992), Powers et al. (1983). Studied localities include Prudhoie Bay and neighbouring North Slope and Brooks Range, as well as localities along Kobuk River near Noorvik and Unalakleet River near Unalakleet.

Fauna of the Russian part of the Arctic lists the largest number of nematode species, including many typical freshwater nematodes, but known from only three distinct geographic regions: Novaya Zemlya archipelago together with Vaygach island, Taymyr peninsula together with Severnaya Zemlya, and the Lena River estuary. So far, only 37 species of freshwater nematodes are know from the Lena River estuary (Gagarin 1995, Gagarin 1999b, Gagarin 2000, Gagarin 2001b). The list of nematodes from the Novaya Zemlya archipelago and Vaygach island includes 81 nematode species, both terrestrial (Steiner 1916b) and freshwater (Gagarin 1997c, Gagarin 1997b, Gagarin 1997a, Gagarin 1999a, Gagarin 2000, Gagarin 2001c, Gagarin 2001b). Arctic areas of the Taymyr peninsula and Severnaya Zemlya are the most intensively studied part of the Arctic with localities including the Taymyr Lake, Enisei River, Putorana Plateau, Cape Chelyuskin – 229 species of freshwater and terrestrial nematodes were reported from different sampling sites of this region over the past 40 years (Elshishka et al. 2012, Elshishka et al. 2013, Gagarin 1990, Gagarin 1991a, Gagarin 1991b, Gagarin 1993, Gagarin 1996, Gagarin 1999a, Gagarin 2001b, Gagarin 2001a, Gagarin 2008, Gagarin 2009, Gagarin and Kuzmin 1972, Gagarin and Kuzmin 1983, Kuzmin 1972, Kuzmin 1976, Kuzmin 1986, Kuzmin and Gagarin 1990).

In summary, 391 species, belonging to 146 genera, 54 families, and 10 orders of the phylum Nematoda are known from the entire Arctic (including species of dubious taxonomic status, but exclusing single nomen nudum). Even taking into consideration relatively low variety of habitats in this region, comparing to other areas of the world, as well as the rather harsh environmental conditions, this list is considered to be very incomplete and preliminary.

 Acknowledgements

This project was in part supported by two grants from the Swedish Taxonomy Initiative: "Taxonomy and distribution of free-living nematodes of the order Plectida in Sweden" and "Systematics of Swedish free-living nematodes of the orders Desmodorida and Araeolaimida".

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Known localities in the Arctic for which detailed information about nematode fauna (identified to species level) is available in the literature.