Biodiversity Data Journal Biodiversity Data Journal Biodiversity Data Journal BDJ 1314-2836 1314-2828 Pensoft Publishers Biodiversity Data Journal 10.3897/BDJ.2.e1116 3155 Taxonomic paper Evanioidea Hymenoptera Faunistics & Distribution Catalogues and Checklists British Isles Europe Checklist of British and Irish Hymenoptera - Evanioidea Broad Gavin R. Dr g.broad@nhm.ac.uk Livermore Laurence Mr The Natural History Museum, London, United Kingdom

Corresponding author: Gavin R. Broad (g.broad@nhm.ac.uk).

Academic editor: Jose Fernandez-Triana.

2014 17 06 2014 2 e1116 20 05 2014 15 06 2014 Gavin R. Broad, Laurence Livermore This is an open access article distributed under the terms of the Creative Commons Attribution License 4.0 (CC-BY), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Introduction

Abbreviations, rationale for species inclusion and additional information are given in the introductory article for the series. This article is the first in a series of 13 checklist sections covering the Hymenoptera of the British Isles. For each genus and species the original combination and a list of synonyms is provided under the heading 'nomenclature'.

A corresponding up-to-date checklist for Evanioidea is maintained on the Hymenoptera of the British Isles site.

Distribution data for the Evanioidea are taken from Crosskey (1951), with additional references given under notes for each taxon. This is a small superfamily of three extant families, but particularly poorly represented in northern Europe. The Evaniidae (sometimes referred to as 'ensign wasps') are, where known, parasitoids/predators of cockroach oothecae (Crosskey 1951; Brown 1973) and are morphologically very distinct from the remaining families. Whilst rather similar in appearance, the Aulacidae and Gasteruptiidae (Fig. 1) differ markedly in biology, with the single British aulacid, Aulacus striatus (Jurine), being a koinobiont endoparasitoid of Xiphydriidae larvae (see E.R. Skinner & W.G. Thompson's 1960 film, 'The Alder Woodwasp and its Insect Enemies', distributed by World Educational Films: http://worldeducationalfilms.com/the-alder-woodwasp-and-its-insect-enemies/) whilst Gasteruption species are essentially predators and cleptoparasites of solitary bee larvae (Malyshev 1964).

Checklists <tp:taxon-name> <tp:taxon-name-part taxon-name-part-type="superfamily">Evanioidea</tp:taxon-name-part> </tp:taxon-name> Aulacidae Shuckard, 1841 Notes

Taxonomy follows Smith (2001).

Aulacus Jurine, 1807 DISPHAERON Dahlbom, 1837 AULACINUS Westwood, 1868 PAMMEGISCHIA Provancher, 1882 DISAULACINUS Kieffer, 1910 MICRAULACINUS Kieffer, 1910 NEURAULACINUS Kieffer, 1910 PARAFOENUS Kieffer, 1910 PYCNAULACUS Cushman, 1929 Aulacus striatus Jurine, 1807 Aulacus arcticus Dahlbom, 1837 Aulacus exaratus Ratzeburg, 1852 Distribution

England.

Evaniidae Latreille, 1802 Notes

Taxonomy follows Deans (2005).

Evania Fabricius, 1775 Evania appendigaster (Linnaeus, 1758) Ichneumon appendigaster Linnaeus, 1758 Ichneumon niger (de Geer, 1773, Ichneumon) Evania laevigata Olivier, 1792 Evania unicolor Say, 1824 Evania desjardinsi Blanchard, 1840 Evania affinis Le Guillou, 1841 Evania cubae Guérin-Méneville, 1844 Evania peringueyi Cameron, 1906 Notes

# Presumably an introduction to Britain as it parasitises the oothecae of non-native synanthropic cockroaches (Blattidae). Crosskey (1951) could find no locality data for British specimens and there are no relevant specimens in NHM.

Brachygaster Leach, 1830 Brachygaster minutus (Olivier, 1792) Evania minuta Olivier, 1792 Evania fulvipes (Curtis, 1829, Evania) Brachygaster rufipes Brullé, 1846 preocc. Evania brullei (Westwood, 1851, Evania) Brachygaster schlettereri Kieffer, 1912 Distribution

England.

Gasteruptiidae Ashmead, 1890 Notes

Synonymic data mostly from Fauna Europaea.

Gasteruption Latreille, 1796 FOENUS Fabricius, 1798 RHYDINOFOENUS Bradley, 1909 DOLICHOFOENUS Kieffer, 1910 TRICHOFOENUS Kieffer, 1910 PLUTOFOENUS Kieffer, 1911 Gasteruption assectator (Linnaeus, 1758) Ichneumon assectator Linnaeus, 1758 Gasteruption affectator misspelling Ichneumon annularis (Geoffrey 1785, Ichneumon) Foenus incertus (Cresson 1864, Foenus) Foenus montanus (Cresson 1864, Foenus) Foenus arca (Couper 1870, Foenus) Foenus borealis (Thomson 1883, Foenus) Foenus fumipennis (Thomson 1883, Foenus) Foenus nigritarsis (Thomson 1883, Foenus) Gasteruption nitidulum Schletterer, 1885 Gasteruption brevicauda Kieffer 1904 Gasteruption micrura Kieffer 1904 Gasteruption nevadense Kieffer 1904 Gasteruption nigropectus Kieffer, 1904 Gasteruption bakeri Kieffer 1910 Trichofoenus canadensis (Kieffer 1910, Trichofoenus) Gasteruption abeillei Kieffer 1912 Gasteruption aberrans Strand, 1912 Trichofoenus breviterebrae (Watanabe 1934, Trichofoenus) Gasteruption utahensis Townes 1950 Gasteruption margotae Madl 1987 Distribution

England, Wales, Ireland.

Gasteruption jaculator (Linnaeus, 1758) Ichneumon jaculator Linnaeus, 1758 Foenus granulithorax (Tournier, 1877, Foenus) Foenus obliteratus (Abeille de Perrin, 1879, Foenus) Foenus rugidorsus (Costa, 1884, Foenus) Gasteruption thomsoni Schletterer, 1885 Distribution

England.

Gasteruption laticeps (Tournier, 1877) Foenus laticeps Tournier, 1877 Gasteruption foveolatum Schletterer, 1889 Gasteruption foveolum Szépligeti, 1903 Gasteruption minutum (Tournier, 1877) Foenus minutum Tournier, 1877 Foenus longigena (Thomson, 1883, Foenus) Distribution

England, Wales (Alexander 1983).

Gasteruption pedemontanum (Tournier, 1877) Foenus pedemontanum Tournier, 1877 Gasteruption pedmontanum misspelling Foenus terrestre (Tournier, 1877, Foenus) Gasteruption trifossulatum Kieffer, 1904 Distribution

England.

References Alexander K. N. A. 1983 Gasteruption minutum (Tournier) (Hymenoptera, Gasteruptiidae) on Anglesey Entomologist's Monthly Magazine 119 150 Brown V. K. 1973 The biology and development of Brachygaster minutus Olivier (Hymenoptera: Evaniidae), a parasite of the oothecae of Ectobius spp. (Dictyoptera: Blattidae) Journal of Natural History 7 6 665 674 http://dx.doi.org/10.1080/00222937300770561 10.1080/00222937300770561 Crosskey R. W. 1951 The morphology, taxonomy, and biology of the British Evanioidea (Hymenoptera) Transactions of the Royal Entomological Society of London 102 247 301 10.1111/j.1365-2311.1951.tb00749.x Deans Andrew R. 2005 Annotated catalog of the world's ensign wasp species (Hymenoptera: Evaniidae) Contributions of the American Entomological Institute 34 1 1 164 Malyshev S. I. 1964 A comparative study of the life and development of primitive gasteruptiids (Hymenoptera, Gasteruptiidae) Entomological Review 43 267 271 Smith David R 2001 World catalog of the family Aulacidae (Hymenoptera) Contributions of the American Entomological Institute 4 263 319

Male Gasteruption jaculator (Linnaeus) (Wilmslow, Cheshire) (courtesy of A.M. Broome).