Corresponding author: Andrey V. Frolov (
Academic editor: Lyubomir Penev
New locality records for
Madagascar houses a rich and taxonomically diverse fauna of the scarab beetles of subfamily
In 2000s, researchers from the California Academy of Sciences (San-Francisco, USA) undertook insect surveys on Madagascar which yielded a rich orphnine material including a few new species (
All examined material is housed in the collection of California Academy of Sciences (San-Francisco, USA).
Preparation of genitalia follows the common technique used in entomological research. The photograph was taken with a Leica MZ9.5 stereo microscope and a Leica DFC290 digital camera from specimen in glycerol. Partially focused serial images were combined in Helicon Focus software (Helicon Soft Ltd.) to produce completely focused image. The photograph was not altered except for levels and tone correction in Adobe Photoshop (Adobe Inc.). The distribution map was generated with ArcGIS software. As the base map, a Madagascar vegetation map (CEPF Madagascar Vegetation Mapping Project of Royal Botanic Gardens, Kew, Missouri Botanical Garden, and Conservation International’s Center for Applied Biodiversity Science;
This species was known from 2 types from Perinet (Andasibe). The two additional localities reported here are situated 70 km NNW and 140 km NNE of the type locality, both within the humid forest biome (Fig.
Type specimens of
This species was known from the only type specimen collected in Antanambe, on the eastern coast of Madagascar. The additional specimen (Fig.
The species was described from one specimen from Ambatofitorahana. The new locality records include Itremo Massif and Ranomafana National Park (Fig.
This species was described from two rather distant localities, in the Itremo Massif and Tsiroanomandidy (Bongolava district). New records extend the known species range to the eastern slopes of the central plateau occupied by the largest remnants of rain forest (Fig.
I am thankful to Dave Kavanaugh, Jere Schweikert, Igor Sokolov, and Norm Penny for help during my visit to California Academy of Sciences. This work was supported by the Ernst Mayr Travel Grant and, partly, by Russian Foundation for Basic Research (grant 13-04-01002-a).